Chamguava: A New Genus of Myrtaceae (Myrtinae) from Mesoamerica. L. R. Landrum. 1991. Systematic Botany

Shrub or tree to 20 m tall, essentially glabrous or puberulent on the disk, petals, and inner calyx surface; trichomes reddish brown to whit­ish, to ca. 0.3 mm long; young twigs gray to reddish brown, remaining more or less smooth, terete or weakly 4-angled. Leaves elliptic to ob­long-elliptic, 3.5-15.5 cm long, 2.2-5 cm wide, 1.6-3.4 times longer than wide; apex acuminate; base rounded, acute or acuminate; petiole 2-4 mm long, 1.5-2 mm thick, shallowly sulcate to flat above; lateral veins rather weak, ca. 14-25 pairs reaching the marginal vein, ascending somewhat, nearly straight; marginal vein about equalling the laterals in prominence, about par­allel to the margin; blades coriaceous to submembranous, drying dark olive-green, gray-green, or reddish brown, somewhat darker above than below. Peduncles 0-2.3 cm long, ca. 0.7 mm wide; bracteoles subtriangular, mem­branous, 1-1.2 mm long and wide, glabrous or with a ciliate margin, persisting until after anthesis; calyx closed or closed except for an apical pore in bud, glabrous without, puberulent or glabrous within, campanulate with an apiculate apex before anthesis, at anthesis tearing irreg­ularly or in 4 lobes, the remnants 3-4 mm long, sometimes persisting until the fruit matures; petals 4-7 mm long, puberulent or glabrous ex­cept for a ciliate margin; hypanthium 2.5-3 mm long, obconic to campanulate, glabrous, densely glandular; disk 4-7 mm across, puberulent on staminal ring; stamens 75-100, 5-6 mm long; anthers ca. 0.8 mm long, the connective with 4-9 glands; style glabrous, 6-7 mm long; ovules (2)10-80 per locule. Fruit subglobose, 1-1.5 cm in diam.; seed 0.8-1 cm long.

Eugenia schippii Standl. Glabrous tree, 10-20 m tall, to 40 cm dbh; stems terete, grayish, the nodes somewhat to conspicuously swollen, the branching mostly dichotomous. Petioles 1-5 mm long, +/- terete or flattened on upper surface, sometimes roughened; blades oblong-elliptic, gradually long-acuminate (the acumen sometimes downturned), broadly to narrowly acute to very weakly attenuate at base, 7-16 (20) cm long, 2-5 cm wide, markedly bicolorous, both surfaces dull, the upper surface with sunken midrib, the lower surface and midrib prominently punctate, the margins weakly revolute; lateral veins 2-6 mm apart, obscure, with a collecting vein 2-3 mm from margin, the reticulate veins very obscure; new flush of leaves +/- maroon. Flowers few to several at leafless, swollen nodes; pedicels 2-5 mm long, the bracts solitary or paired, borne apically, +/- round or ovate to +/- acute, pellucid-punctate, ciliate, usually closely appressed to the hypanthium; flower buds globose to obovoid, ca 7 mm long; calyx completely closed in bud, splitting +/- irregularly, 2-5-lobed, the lobes rounded to acute at apex, 3-3.5 mm long, persisting in fruit; petals 5, +/- rounded, concave, inconspicuously ciliate, white, ca 5 mm long; stamens numerous; stamens and style +/- equaling petals; style glabrous, narrowly tapered to apex; ovules numerous, flattened and reniform, less than 1 mm diam. Berries globose to depressed-globose, 2-3.5 cm diam, green tinged with red (probably also becoming completely red) when mature; seeds usually 2, horseshoe-shaped, ca 2 cm long and 3 mm wide. Croat 16213. Apparently rare, in the old forest; known from the vicinity of Zetek Trail 300-400, where seedlings are common. Mature fruits have been seen on BCI in July. Both flowers and immature fruits are seen elsewhere in Panama in August. In Belize the flowers are seen in May and mature fruits in August and November. Owing to its unusual C-shaped embryo, which lacks a hard bony testa, it is possible that the species does not even belong in the genus Psidium (R. McVaugh, pers. comm.). Panamanian material differs from that of plants from Belize in having shorter pedicels and calyx lobes that are essentially glabrous inside rather than strongly pubescent. Fruits on Lao & Holdridge 194 (Salúd, Colón Province) were heavily infested with gall-forming insects, which caused the fruits to be somewhat lobed.